Are they comfortable to the guys that used to wear old Linds, probably. Are they comfortable to the generation that thinks flip flops are shoes, maybe not.
My take-The two inches of hard plastic at the front of the shoe that doesn't flex definitely feels different and takes some getting used to, and for a shoe that is going to retail for $190 Dexter could have sprung for a little better insole.
I'm staring at the shoe as I write this, and the only hard plastic I see is the reenforcement at the tip where the replaceable sole with the toe guard slots in. It's not a part of the shoe you would normally expect to flex, unless you're doing some kind of ballerina pose at the finish. I'm trying to understand where there should be any 'feel' difference.
As far as insoles, for most any shoe (of any type) made, the generic footbeds are crap. I don't blame the manufacturers because they don't know the shape of the foot going into the shoe. The person could have a high arch, no arch, or anything in-between. You try to cater to one, and every other foot type is going to be extra miserable.
That's why for almost every shoe I own, I insert an orthodic that conforms to the shape and support needs of my feet. That means my ski boots, snowboarding boots, hard hiking boots, soft hiking boots, tennis shoes, casual walking shoes, and of course, bowling shoes. I'm still getting the feel for my 9s, and haven't put in my favorite orthodic yet. I'm surprised that so far, I haven't felt the need to.