After looking at some videos (Don't have a very big library, only a few bowlers). It would seem that a bowler would be using the side (or corner) of their fingers to put revs on the ball to get more tilt.
Take the extreme example. Spinner Release (Very High Tilt). They are clearly not revving the ball with the pads of their fingers, instead they come around the ball.
Another example. I know some bowlers who use grips and they are constantly wearing out the side of their finger grips. Obviously these bowlers are using the side of their fingers to get revs which probably increases their tilt.
This is up for debate. Just throwing my ideas out there.